Brontosaurus is a genus of herbivorous sauropod dinosaur that lived in present-day United States during the Late Jurassic period. It was described by American paleontologist Othniel Charles Marsh in 1879, the type species being dubbed B. excelsus, based on a partial skeleton lacking a skull found in Como Bluff, Wyoming. In subsequent years, two more species of Brontosaurus were named: B. parvus in 1902 and B. yahnahpin in 1994. Brontosaurus lived about 156 to 146 million years ago during the Kimmeridgian and Tithonian ages in the Morrison Formation of what is now Utah and Wyoming. For decades, the animal was thought to have been a taxonomic synonym of its close relative Apatosaurus, but a 2015 study by Emmanuel Tschopp and colleagues found it to be distinct. It has seen widespread representation in popular culture, being the archetypal "long-necked" dinosaur in general media.